Bees might learn that the colors of petals, anthers, or pollen itself predicts pollen rewards.
In our new Animal Behaviour paper (Muth et al. 2016 or try ResearchGate) , Felicity pushed the study of pollen learning forward by 1) demonstrating that bees can learn based on reinforcement by floral-collected pollen and 2) exploring what features of flowers they can learn, and how long they remember these associations.